Editor(s):
D. Geltman & R. Riina


  Name:      Euphorbia virgata   Waldst. & Kit., Descr. Icon. Pl. Hung. 2: 176. 1805 publ. 1803.

Status:accepted_name
Accepted Name:None
Basionym:None
Common Names:
Leafy spurge, wolf's milk
TOLKIN GUID:72e402f9-ed6d-4c86-aaaa-a3068d7d9478
TOLKIN GUID URL:http://app.tolkin.org/guids/72e402f9-ed6d-4c86-aaaa-a3068d7d9478
Type Details:
Species: None
Type Collection:
Waldstein s.n.
Type Date: None
Type Locality:
Si altiora loca montosa demas, per omnem fere Hungariam, Croatiam, & Sclavoniam locis siccis.
Type Herbaria:
PR, photo E
Neo Type: None
Synonyms: show/hide
Name
Namestatus
Tithymalus boissierianus Woronow, Herb. Fl. Cauc. 479. 1931.
Synonym
Tithymalus hypoleucus Prokh., Consp. Syst. Tithymalus As. Med. 199. 1933.
Synonym
Euphorbia hypoleuca (Prokh.) Rech., Ann. Naturhist. Mus. Wien 56: 212. 1948.
Synonym
Euphorbia boissieriana (Woronow) Prokh., Fl. URSS 14: 445. 1949.
Synonym
Euphorbia minxianensis W.T. Wang, Acta Bot. Yunnan. 10: 43. 1988.
Synonym
Subgenus:
Esula
Section:
Esula
Subsection: None
Ingroup Clade:
B
Description:
Perennial herbs, from a strong branching rootstock. Stems erect, simple or branched, glabrous, terete, 20-90 cm. Leaves alternate; petiole absent or short, 0.5-1 mm, glabrous; blade linear to linear-oblanceolate or linear-oblong (margins parallel or almost parallel at mid-leaf), 40-90 x 3-12 mm, venation inconspicuous, only mid-vein prominent, glabrous, base truncate or adruptly attenuate, apex acute or rounded, sometimes mucronulate, margin entire. Cyathia in leafy pleiochasial rays; peduncle absent or very short (less than 1 mm); terminal rays 5-17, 1-2 times bifurcated; ray-leaves similar but shorter and wider than the upper stem leaves; subcyathial raylet-leaves, widely ovate, rhombic or reniform, base cordate or cuneate, apex obtuse to rounded, mucronulate, margin entire; axillary rays 0-18. Involucre campanulate, 1.5-3.5 x 1.7-3 mm, glabrous; glands 4, crescent-shaped, 0.6-1.5 x 1.3-2.5 mm; horns present, divergent to convergent, 0.2-0.8 mm. Staminate flowers 10-25. Pistillate flower: ovary glabrous, styles 1.7-2.5 mm, bifid. Capsule subglobose, slightly lobed, cocci rounded, smooth, finely granular toward the dorsal line, glabrous, 2.5-3.5 x 3-4.5 mm; columella 2-3.3 mm. Seeds oblong-elliptic to oblong-ovoid, 2.2—2.6 x 1.3-1.6 mm, smooth, yellow-brown to gray or mottled; caruncle subconical, 0.6-1 x 0.7-0.9 mm.

Flowering and fruiting spring to fall.
Habitat:
Pastures, fields and waste places, railroads, open disturbed areas; 0--2200 m.
Phylogenetic Relationships: None
Comments:
Native to Europe and temperate Asia and introduced to North America. We are treating this species here as the widespread, weedy, “leafy spurge” that is a pest plant across much of the northern parts of the FNA area. This species is similarly widespread and weedy across much of its native range in Europe and Asia, whereas the true E. esula is much more restricted in range and is never as pervasive and weedy as E. virgata in either its native range or the FNA area.
Chromosome Number: None
Uses: None
Toxicity: None
Conservation Status: None
General Distribution:
Central Europe to Causasus and South-West Asia: Denmark, Austria, Czech Republic, Slovakia, Germany, Hungary, Switzerland, Spain, Bulgaria, Italy, Romania, Ex-Yugoslavia, Estonia, Kaliningrad, Lithuania, Latvia, Russia, Belarus, Moldova, Ukraine, Crimea, Kazakhstan, Iran, Iraq, Turkey. Introduced to North America; Canada (British Columbia, Ontario, Quebec); U.S.A. (Alaska, Arizona, California, Colorado, Connecticut, Idaho, Indiana, Illinois, Iowa, Kansas, Maine, Maryland, Massachusetts, Michigan, Minnesota, Missouri, Montana, Nebraska, Nevada, New Hampshire, New Jersey, New Mexico, New York, North Dakota, Oregon, Pennsylvania, South Dakota, Utah, Virginia, Vermont, Wisconsin, Washington, Wyoming).
Collector
Collection number
Country
Institution code
Frajman, B.
11101
Austria
Pahlevani, A.
53772
Iran
IRAN
Berry, Paul E.
8016
Russia
MICH
Berry, Paul E.
8017
Russia
MICH
Berry, Paul E.
8018
Russia
LE, MICH
Berry, Paul E.
8020
Russia
LE, MICH
Berry, Paul E.
8021
Russia
LE, MICH
Berry, Paul E.
8022
Russia
LE
Riina, R.
1868
Spain
MA
Riina, R.
1868
Spain
BCN
DNA Sample(s): 8846
DNA Sample(s) Sequences
References:

Geltman, D.V. (1998). Taxonomic notes on Euphorbia esula (Euphorbiaceae) with special reference to its occurrence in the east part of the Baltic region. Ann. Bot. Fennici 35, 113-117.

Govaerts, R., Frodin, D.G., and Radcliff-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae). The Board of Trustees of the Royal Botanic Gardens, Kew.

Riina, R., Peirson, J.A., Geltman, D.V., Molero, J., Frajman, B., Pahlevani, A., Barres, L., Morawetz, J.J., Salmaki, Y., Kryukov, A., Bruyns, P.V., and Berry, P.E. (2013). A worldwide molecular phylogeny and classification of the leafy spurges, Euphorbia subgenus Esula (Euphorbiaceae). Taxon 62, 316–342.


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